• Associate Professor
  School of Natural Sciences and Engineering
  Ilia State University
  3/5 Choloqashvili Ave
  Tbilisi 0162
  Republic of Georgia
• ORCID
• Ongoing Habilitation (Professorial Dissertation): Ecology, Biogeography and DNA Barcoding in Athropods of Alpine Tundra Ecosystems
• Biocoenology, Biogeography, DNA Barcoding, Arthropods, Araneae, Alpine Tundra Ecosystems
• Norway, Georgia
• Co-Chair of German Biogeography Study Group (AK Biogeographie, DGfG)

Personal webpage LIB

ResearchGate

Articles in peer-reviewed journals

2024

RDUCH, V., DEY, L.-S., GHAZARYAN, A., HUSEMANN, M., IANKOSHVILI, G., KRAMMER, H.-J., LAMBRECHT, M., MARABUTO, E., MENGUAL, X., MÜLLER, B., PETERS, R.S., RULIK, B., SALDEN, T., SILVA-BRANDÃO, K.L., ESPELAND, M., THORMANN, J., TÖPFER, T., WEISS, C., ŻYŁA, D. & HEIN, N (2024) Exkursion in den Kaukasus: Der CaBOL BioBlitz 2022 in Armenien und Georgien. Koenigiana, 18, 21-40.

SALDEN, T., MÜLLER, B., JAPOSHVILI, G., HEIN, N., UGRELIDZE, A. & PETERS, R.S. (2024) First records of the Hymenoptera superfamilies and families Mymarommatoidea: Mymarommatidae and Stephanoidea: Stephanidae in Georgia. Caucasiana, 3, 145-150. https://doi.org/10.3897/caucasiana.3.e124925

WELLENBECK, A., FEHRMANN, L., FEILHAUER, H., SCHMIDTLEIN, S., MISOF, B., & HEIN, N. (2024). Discriminating Woody Species Assemblages from National Forest Inventory Data Based on Phylogeny in Georgia. Ecology and Evolution, 14, e11569. https://doi.org/10.1002/ece3.11569

HEIN, N., ASTRIN, J., BECKERS, N., GIEBNER, H., LANGEN, K., LÖFFLER, J., MISOF, B. & FONSECA, V. (2024) Arthropod Diversity in the Alpine Tundra using Metabarcoding: Spatial and Temporal Differences in Alpha- and Beta-Diversity. Ecology and Evolution. http://dx.doi.org/10.1002/ece3.10969

TARKHNISHVILI, D., SEROPIAN, A., ERHARDT, C., KACHLISHVILI, N., KRAMMER, H.-J., & HEIN, N. (2024) How Dispersal Rates Depend on the Prey Capture Strategy: A Case Study of Georgia´s Spiders. Ecology and Evolution, 14, e11372. https://doi.org/10.1002/ece3.11372

SEROPIAN, A., BULBULASHVILI, N., KRAMMER, H.-J., THORMANN, J., HEIN, N., KARALASHVILI, E., KACHLISHVILI, N. & DATUNASHVILI, A. (2024) Picking Pearls from the Silk Road: Insights into the Spider (Arachnida, Araneae) Diversity in Georgia from the Caucasus Barcode of Life (CaBOL) Project. Part III. Caucasiana, 3, 89-118. https://doi.org/10.3897/caucasiana.3.e120883

HEIN, N., ASTRIN, J., BECKERS, N., GIEBNER, H., LANGEN, K., LÖFFLER, J., MISOF, B. & FONSECA, V. (2024) Arthropod Diversity in the Alpine Tundra using Metabarcoding: Spatial and Temporal Differences in Alpha- and Beta-Diversity. Ecology and Evolution. http://dx.doi.org/10.1002/ece3.10969

2023

SEROPIAN, A., ARSENASHVILI, E., BULBULASHVILI, N., SHUBASHISHVILI, A., IANKOSHVILI, G., TODUA, M., ANANIASHVILI, A., JAPARASHVILI, S., CHKHARTISVHILI, T., MEMISHISHI, A., BALKHAMISHVILI, S., CHITADZE, B., KARALASHVILI, E., MUMLADZE, L., HEIN, N. & RULIK, B. (2023) Into the Unknown: the First Barcode-Assisted Checklist of Psocoptera (Insecta, Psocodea) of Georgia with a Census on Country Species Richness. ZooKeys, 1168, 77.

WEHNER, A., HEIN, N., DOBBERT, S., PAPE, R., LÖFFLER, J. (2023) Early Snow Melt and Diverging Thermal Constraints Control Body Size in Arctic-Alpine Apiders. Biological Journal of the Linnean Society, 138, 1–13. https://doi.org/10.1093/biolinnean/blac127

2022

WEHNER, A., HEIN N., BECKERS, N., DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) Data from: Early Snow Melt and Diverging Thermal Constraints Control Body Size in Arctic-Alpine Spiders. Dryad Digital Repository. https://doi.org/10.5061/dryad.18931zd1c

2021

ANTHONY, S. E., BUDDLE, C. M., HØYE, T. T., HEIN, N. & SINCLAIR, B. J. (2021) Thermal Acclimation has Limited Effect on the Thermal Tolerances of Summer-Collected Arctic and Sub-Arctic Wolf Spiders. Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, 257, 110974. https://doi.org/10.1016/j.cbpa.2021.110974

HEIN, N., MERKELBACH, J., ZECH, K. & WEIJERS, S. (2021) Drought Sensitivity of Empetrum nigrum Shrub Growth at the Species' Southern Lowland Distribution Range Margin. Plant Ecology, 222, 305–321. https://doi.org/10.1007/s11258-020-01107-z

2020

KANKAANPÄÄ, T., VESTERINEN, E., HARDWICK, B. et al. (2020) Parasitoids Indicate Major Climate-Induced Shifts in Arctic Communities. Global Change Biology, 00, 1–20. https://doi.org/10.1111/gcb.15297

BECKERS, N., HEIN, N., ANNESER, A., VANSELOW, K. A. & LÖFFLER, J. (2020) Differences in Mobility and Dispersal Capacity Determine Body Size Clines in Two Common Alpine-Tundra Arthropods. Insects, 11 (2), 74. https://doi.org/10.3390/insects11020074

2019

HEIN, N., LÖFFLER, J. & FEILHAUER, H. (2019) Mapping of Arthropod Alpha and Beta Diversity in Heterogeneous Arctic-Alpine Ecosystems. Ecological Informatics, 54, 101007. https://doi.org/10.1016/j.ecoinf.2019.101007

HEIN, N., PÉTILLON, J., PAPE, R., FEILHAUER, H., VANSELOW, K. A. & LÖFFLER, J. (2019) Broad-Scale Rather than Fine-Scale Environmental Variation Drives Body Size in a Wandering Predator (Araneae, Lycosidae). Arctic, Antarctic, and Alpine Research, 51, 315–326. https://doi.org/10.1080/15230430.2019.1640039

2018

BECKERS, N., HEIN, N., VANSELOW, K. A. & LÖFFLER, J. (2018) Effects of Microclimatic Thresholds on the Activity-Abundance and Distribution Patterns of Alpine Carabidae Species. Annales Zoologici Fennici, 55, 25–44. https://doi.org/10.5735/086.055.0104

HEIN, N., BRENDEL, M. R., FEILHAUER, H., FINCH, O.-D. & LÖFFLER, J. (2018) Egg Size vs. Egg Number Trade-Off in the Alpine-Tundra Wolf Spider, Pardosa palustris (Araneae: Lycosidae). Polar Biology, 41, 1607–1617. https://doi.org/10.1007/s00300-018-2301-x

2015

HEIN, N., FEILHAUER, H., LÖFFLER, J. & FINCH, O.-D. (2015) Elevational Variation of Reproductive Traits in five Pardosa (Lycosidae) Species. Arctic, Antarctic, and Alpine Research, 47, 67–73. https://doi.org/10.1657/AAAR0013-111

2014

HEIN, N., FEILHAUER, H., FINCH, O.-D., SCHMIDTLEIN, S. & LÖFFLER, J. (2014) Snow Cover Determines the Ecology and Biogeography of Spiders (Araneae) in Alpine Tundra Ecosystems. Erdkunde, 68, 157–172. https://doi.org/10.3112/erdkunde.2014.03.01

HEIN, N., PAPE, R., FINCH, O.-D. & LÖFFLER, J. (2014) Alpine Activity Patterns of Mitopus morio (Fabricius, 1779) are induced by Variations in Temperature and Humidity at Different Scales in Central Norway. Journal of Mountain Science, 11, 644–655. https://doi.org/10.1007/s11629-013-2913-0

2013

HEIN, N., SOLHØY, T., SCHATZ, H. & LÖFFLER, J. (2013) An Oribatid Species Provertex kuehnelti Mihelčič, 1959 (Acari, Oribatida) New to Fennoscandia. Norwegian Journal of Entomology, 60, 163–168.

HÅGVAR, S., HEIN, N., LÖFFLER, J., LUNDMO, S. & STRAUMFORS, P. (2013) New Data on the Distribution of Chionea Dalman, 1816 (Diptera, Limoniidae), and on Habitat Choice of C. araneoides Dalman, 1816 at High Altitudes. Norwegian Journal of Entomology, 60, 176–181.

Other publications

2019

THORMANN, J., AHRENS, D., ANDERSON, C., ASTRIN, J., MUMLADZE, L., RULIK, B., TARKHNISHVILI, D., ESPELAND, M., GEIGER, M., HEIN, N., IANKOSHVILI, G., KARALASHVILI, E., MENGUAL, X., MORKEL, C., NEIBER, M.T., PETERS, R.S., REIMANN, A., SSYMANK, A., WESENER, T., ZIEGLER, J. & MISOF, B. (2019) A prelude to the Caucasus Barcode of Life Platform (CaBOL): Biodiversity Days in Georgia in 2018 and 2019. Bonn Zoological Bulletin. https://doi.org/10.20363/bzb-2019.68.2.275

2015

BECKERS, N., N. HEIN & J. LÖFFLER (2015) Klimaökologische Diversitätsmuster von Laufkäfern (Carabidae) im norwegischen Hochgebirge. Norden, 21, 45-53. Bremen.

2007

BUCHHOLZ, S. & HEIN, N. (2007) Die epigäische Webspinnenfauna (Araneae) verschiedener Habitattypen der Emsaue bei Münster (NRW). Natur und Heimat, 67, 109-124.